Health Sciences and Technology (HST)
IMPORTANT NOTES regarding preclinical subjects (HST.011-HST.200)*:
Students not enrolled in an HST program are limited to two HST preclinical courses and must provide justification for enrolling in these courses. This action must be approved by the course director and the student's advisor. These subjects are scheduled according to the Harvard Medical School academic calendar, which differs from the MIT calendar. Students whose graduation depends upon completing one or more of these subjects should take particular care regarding the schedule. * HST.163 and HST.198 are NOT included in the two-course limit.
HST.011 Human Functional Anatomy
Subject meets with HST.010
Prereq: Permission of instructor
G (Fall)
3-11-10 units
Lectures, detailed laboratory dissections, and prosections provide a thorough exploration of the gross structure and function of the human body. Fundamental principles of bioengineering are employed to promote analytical approaches to understanding the body's design. The embryology of major organ systems is presented, together with certain references to phylogenetic development, as a basis for comprehending anatomical complexity. Correlation clinics stress both normal and abnormal functions of the body and present evolving knowledge of genes responsible for normal and abnormal anatomy. Lecturers focus on current problems in organ system research. Only HST students may register under HST.010, graded P/D/F. Lab fee. Enrollment restricted to graduate students.
T. Van Houten, R. Mitchell
HST.015 MATLAB for Medicine
Prereq: None
G (Summer)
Not offered regularly; consult department
2-0-4 units
Practical introduction to use of quantitative methods in medicine and health research. Each session covers a different topic in quantitative techniques, provides an application to medicine, and includes a modeling activity using MATLAB. Students also complete problem sets. Restricted to first year HST MD students.
M. Frosch
HST.021 Musculoskeletal Pathophysiology
Subject meets with HST.020
Prereq: HST.030 and HST.160
G (Spring)
3-0-3 units
Growth and development of normal bone and joints, the biophysics of bone and response to stress and fracture, calcium and phosphate homeostasis and regulation by parathyroid hormone and vitamin D, and the pathogenesis of metabolic bone diseases and disease of connective tissue, joints, and muscles, with consideration of possible mechanisms and underlying metabolic derangements. Only HST students may register under HST.020, graded P/D/F. Enrollment limited; restricted to medical and graduate students.
M. Bouxsein, L. Tarter
HST.031 Human Pathology
Subject meets with HST.030
Prereq: Biology (GIR), Physics I (GIR), and permission of instructor
G (Fall)
4-3-8 units
Introduction to the functional structure of normal cells and tissues, pathologic principles of cellular adaptation and injury, inflammation, circulatory disorders, immunologic injury, infection, genetic disorders, and neoplasia in humans. Lectures, conferences emphasizing clinical correlations and contemporary experimental biology. Laboratories with examination of microscopic and gross specimens, and autopsy case studies emphasizing modern pathology practice. Only HST students may register under HST.030, graded P/D/F. Lab fee. Enrollment limited.
R. N. Mitchell, R. Padera
HST.041 Mechanisms of Microbial Pathogenesis
Subject meets with HST.040
Prereq: Biology (GIR), 7.05, and permission of instructor
G (Fall)
4-2-6 units
Deals with the mechanisms of pathogenesis of bacteria, viruses, and other microorganisms. Approach spans mechanisms from molecular to clinical aspects of disease. Topics selected for intrinsic interest and cover the demonstrated spectrum of pathophysiologic mechanisms. Only HST students may register under HST.040, graded P/D/F. Lab fee. Enrollment limited.
S. Kanjilal, K. Hysell
HST.061 Endocrinology
Prereq: Biology (GIR), 7.05, and permission of instructor
G (Spring)
3-0-6 units
Physiology and pathophysiology of the human endocrine system. Three hours of lecture and section each week concern individual parts of the endocrine system. Topics include assay techniques, physiological integration, etc. At frequent clinic sessions, patients are presented who demonstrate clinical problems considered in the didactic lectures. Only HST students may register under HST.060, graded P/D/F. Enrollment limited.
W. Kettyle, Y-M. Chan, A. Abreu
HST.071 Human Reproductive Biology
Subject meets with HST.070
Prereq: 7.05 and permission of instructor
G (Fall; first half of term)
4-0-2 units
Lectures and clinical case discussions designed to provide the student with a clear understanding of the physiology, endocrinology, and pathology of human reproduction. Emphasis is on the role of technology in reproductive science. Suggestions for future research contributions in the field are probed. Students become involved in the wider aspects of reproduction, such as prenatal diagnosis, in vitro fertilization, abortion, menopause, contraception and ethics relation to reproductive science. Only HST students may register under HST.070, graded P/D/F.
A. Koniaris, D. Page, T. Lau
HST.081 Hematology
Subject meets with HST.080
Prereq: 7.05 and permission of instructor
G (Spring; partial term)
2-1-3 units
Intensive survey of the biology, physiology and pathophysiology of blood with systematic consideration of hematopoiesis, white blood cells, red blood cells, platelets, coagulation, plasma proteins, and hematologic malignancies. Emphasis given equally to didactic discussion and analysis of clinical problems. Enrollment limited.
D. Bauer, S. Schulman
HST.091 Cardiovascular Pathophysiology
Subject meets with HST.090
Prereq: (HST.030 or HST.031) and permission of instructor
G (Spring)
4-3-8 units
Normal and pathologic physiology of the heart and vascular system. Emphasis includes hemodynamics, electrophysiology, gross pathology, and clinical correlates of cardiovascular function in normal and in a variety of disease states. Special attention given to congenital, rheumatic, valvular heart disease and cardiomyopathy. Only HST students may register under HST.090, graded P/D/F. Enrollment limited.
C. Stultz, T. Heldt
HST.101 Respiratory Pathophysiology
Subject meets with HST.100
Prereq: Physics I (GIR), 7.05, and permission of instructor
G (Spring)
4-0-8 units
Lectures, seminars, and laboratories cover the histology, cell biology, and physiological function of the lung with multiple examples related to common diseases of the lung. A quantitative approach to the physics of gases, respiratory mechanics, and gas exchange is provided to explain pathological mechanisms. Use of medical ventilators is discussed in lecture and in laboratory experiences. For MD candidates and other students with background in science. Only HST students may register under HST.100, graded P/D/F. Enrollment limited.
C. Hardin, E. Roche, K. Hibbert
HST.111 Renal Pathophysiology
Subject meets with HST.110
Prereq: 7.05 and permission of instructor
G (Spring)
4-0-8 units
Considers the normal physiology of the kidney and the pathophysiology of renal disease. Renal regulation of sodium, potassium, acid, and water balance are emphasized as are the mechanism and consequences of renal failure. Included also are the pathology and pathophysiology of clinical renal disorders such as acute and chronic glomerulonephritis, pyelonephritis, and vascular disease. New molecular insights into transporter mutations and renal disease are discussed. Only HST students may register under HST.110, graded P/D/F. Enrollment limited.
G. McMahon, M. Yeung
HST.121 Gastroenterology
Subject meets with HST.120
Prereq: Biology (GIR), Physics I (GIR), 7.05, and permission of instructor
G (Fall; second half of term)
3-1-2 units
Presents the anatomy, physiology, biochemistry, biophysics, and bioengineering of the gastrointestinal tract and associated pancreatic, liver, and biliary systems. Emphasis on the molecular and pathophysiological basis of disease where known. Covers gross and microscopic pathology and clinical aspects. Formal lectures given by core faculty, with some guest lectures by local experts. Selected seminars conducted by students with supervision of faculty. Only HST students may register under HST.120, graded P/D/F. Enrollment limited.
A. Rutherford, S. Flier
HST.131 Neuroscience
Subject meets with HST.130
Prereq: Permission of instructor
G (Fall)
6-3-6 units
Comprehensive study of neuroscience where students explore the brain on levels ranging from molecules and cells through neural systems, perception, memory, and behavior. Includes some aspects of clinical neuroscience, within neuropharmacology, pathophysiology, and neurology. Lectures supplemented by conferences and labs. Labs review neuroanatomy at the gross and microscopic levels. Only HST students may register under HST.130, graded P/D/F. Limited to 50.
J. Assad, M. Frosch
HST.147 Biochemistry and Metabolism
Subject meets with HST.146
Prereq: Permission of instructor
G (Fall)
4-0-5 units
First-year graduate level intensive subject in human biochemistry and physiological chemistry that focuses on intermediary metabolism, structures of key intermediates and enzymes important in human disease. Subject is divided into four areas: carbohydrates, lipids, amino acids and nucleic acids. The importance of these areas is underscored with examples from diseases and clinical correlations. Preparatory sessions meet in August. Only HST students may register under HST.146, graded P/D/F. Enrollment limited.
R. Sharma
HST.151 Principles of Pharmacology
Subject meets with HST.150
Prereq: Biology (GIR), Physics I (GIR), and 7.05
G (IAP, Spring; partial term)
6-0-6 units
An introduction to pharmacology. Topics include mechanisms of drug action, dose-response relations, pharmacokinetics, drug delivery systems, drug metabolism, toxicity of pharmacological agents, drug interactions, and substance abuse. Selected agents and classes of agents examined in detail. Course follows HMS calendar. Restricted to HST MD & HST PhD students.
S. Forman
HST.161 Genetics in Modern Medicine
Subject meets with HST.160
Prereq: 7.05
G (Fall; second half of term)
2-0-4 units
Provides a foundation for understanding the relationship between molecular biology, genetics, and medicine. Starts with an introduction to molecular genetics, and quickly transitions to the genetic basis of diseases, including chromosomal, mitochondrial and epigenetic disease. Translation of clinical understanding into analysis at the level of the gene, chromosome, and molecule; the concepts and techniques of molecular biology and genomics; and the strategies and methods of genetic analysis. Includes diagnostics (prenatal and adult), cancer genetics, and the development of genetic therapies (RNA, viral, and genome editing). The clinical relevance of these areas is underscored with patient presentations. Only HST students may register under HST.160, graded P/D/F.
S. Nissim, R. Gupta
HST.163 Molecular Diagnostics and Bioinformatics
Subject meets with HST.162
Prereq: Permission of instructor
G (Fall; first half of term)
2-0-4 units
Introduction of molecular diagnostic methods in medicine and relevant bioinformatics methods. Discussion of principles of molecular testing for diagnosis of somatic and germline diseases using FISH, classical genotyping, array CGH, next generation sequencing, and other technologies. Case conferences emphasize clinical correlation and integration of information from multiple diagnostic tests. Bioinformatics lectures, problem sets, and laboratory sessions will introduce key concepts in biological sequence analysis and provide experience with bioinformatics tools. HST.015 and HST.191 recommended. Only HST students may register under HST.162, P/D/F. Enrollment limited, preference to HST students.
G. Gerber, L. Li
HST.165 Principles of Biomedical Imaging
Subject meets with HST.164
Prereq: Permission of instructor
Acad Year 2024-2025: Not offered
Acad Year 2025-2026: G (IAP)
2-0-4 units
Reviews fundamental principles and techniques underlying modern biomedical imaging, as well as their application in modern medicine. Particular emphasis on magnetic resonance; also covers ultrasound, computed tomography, positron emission tomography and optical techniques. Didactic lectures accompanied by problem sets and experiments with portable magnetic resonance systems and ultrasound systems. Focuses on the quantitative aspects of biomedical imaging and requires a knowledge of differential equations, MATLAB, and intermediate-level physics. Only HST students may register under HST.164, P/D/F. Restricted to HST students.
S. Huang, D. Sosnovik
HST.176 Cellular and Molecular Immunology
Subject meets with HST.175
Prereq: 7.05
G (Fall)
6-0-6 units
Covers cells and tissues of the immune system, lymphocyte development, the structure and function of antigen receptors, the cell biology of antigen processing and presentation including molecular structure and assembly of MHC molecules, lymphocyte activation, the biology of cytokines, leukocyte-endothelial interactions, and the pathogenesis of immunologically mediated diseases. Consists of lectures and tutorials in which clinical cases are discussed with faculty tutors. Details of each case covering a number of immunological issues in the context of disease are posted on a student website. Only HST students may register under HST.175, graded P/D/F. Limited to 45.
S. Pillai, B. Cherayil
HST.191 Introduction to Biostatistics
Subject meets with HST.190
Prereq: Calculus II (GIR)
G (Summer)
3-0-3 units
Provides training on how to comprehend, critique and communicate findings from biomedical literature. Considers how to assess the importance of chance in the interpretation of experimental data. Topics include probability theory, chi-squared and t-tests, ANOVA, linear and logistic regression, survival analysis, and statistical analysis using MATLAB. Includes critical reading of studies published in medical literature. Enrollment limited; restricted to HST students.
N. Hejazi
HST.192 Medical Decision Analysis and Probabilistic Medical Inference
Prereq: Permission of instructor
Acad Year 2024-2025: Not offered
Acad Year 2025-2026: G (IAP)
2-0-2 units
Teaches the essentials of quantitative diagnostic reasoning and medical decision analysis. Guides participants through the process of choosing an appropriate contemporary medical problem in which risk-benefit tradeoffs play a prominent role, conducting a decision analysis, and ultimately publishing the results in a medical journal. Topics include decision trees, influence diagrams, Markov decision models and Monte Carlo simulation, methods for quantifying patient values, Bayesian inference, decision thresholds, and the cognitive science of medical decision making. HST.191 recommended. Limited to 8; preference to HST students.
M. B. Westover
HST.195 Clinical Epidemiology
Subject meets with HST.194
Prereq: HST.190
G (IAP, Spring; first half of term)
1-0-1 units
Introduces methods for the generation, analysis, and interpretation of data for clinical research. Major topics include the design of surveys, predictive models, randomized trials, clinical cohorts, and analyses of electronic health records. Prepares students to formulate well-defined research questions, design data collection, evaluate algorithms for clinical prediction, design studies for causal inference, and identify and prevent biases in clinical research. Emphasizes critical thinking and practical applications, including daily assignments based on articles published in major clinical journals and the discussion of a case study each week. Trains students to comprehend, critique, and communicate findings from the biomedical literature. Familiarity with regression modeling and basic statistical theory is a prerequisite. Only HST students may register under HST.194, graded P/D/F. Enrollment limited; restricted to medical and graduate students.
M. Hernan
HST.196 Teaching Health Sciences and Technology
Prereq: None
G (Fall, IAP, Spring, Summer)
Units arranged [P/D/F]
Can be repeated for credit.
Provides teaching experience (classroom, laboratory, field, recitation, tutorial) under the direction of faculty member(s). Students may prepare instructional materials, lead discussion groups, provide individualized instruction, monitor students' progress, and gain experience delivering other educational elements. Limited to qualified graduate students.
HST Faculty
HST.198 Independent Study in Health Sciences and Technology
Prereq: Permission of instructor
G (Fall, IAP, Spring, Summer)
Units arranged
Can be repeated for credit.
Opportunity for independent study of health sciences and technology under regular supervision by an HST faculty member. Projects require prior approval from the HST Academic Office, as well as a substantive paper.
HST Faculty
HST.200 Introduction to Clinical Medicine
Prereq: Permission of instructor
G (IAP, Spring; partial term)
9-19-12 units
Intensive preparation for clinical clerkships that introduces the basic skills involved in examination of the patient in addition to history taking and the patient interview. Provides exposure to clinical problems in medicine, surgery, and pediatrics. Students report their findings through history taking and oral presentations. Restricted to MD program students.
W. Goessling, D. Rubinson, D. Solomon
HST.201 Introduction to Clinical Medicine and Medical Engineering I
Prereq: Permission of instructor
G (Summer)
0-20-0 units
Develop skills in patient interviewing and physical examination; become proficient at organizing and communicating clinical information in both written and oral forms; begin integrating history, physical, and laboratory data with pathophysiologic principles; and become familiar with the clinical decision-making process and broad economic, ethical, and sociological issues involved in patient care. There are two sections: one at Mount Auburn Hospital and one at West Roxbury VA Hospital, subsequent registration into HST.202 must be continued at the same hospital as HST.201. Restricted to MEMP students.
C. Stultz, N. Price, J. Strymish
HST.202 Introduction to Clinical Medicine and Medical Engineering II
Prereq: HST.201
G (Fall, IAP, Spring, Summer)
0-20-0 units
Strengthens the skills developed in HST.201 through a six-week clerkship in medicine at a Harvard-affiliated teaching hospital. Students serve as full-time members of a ward team and participate in longitudinal patient care. In addition, students participate in regularly scheduled teaching conferences focused on principles of patient management. Restricted to MEMP students.
C. Stultz, N. Price, J. Strymish, A. Romano
HST.207 Introduction to Clinical Medicine and Medical Engineering
Prereq: Permission of instructor
G (Spring; partial term)
0-30-0 units
Introduction to the intricacies of clinical decision-making through broad exposure to how clinicians think and work in teams. Instruction provided in patient interviewing and physical examination; organizing and communicating clinical information in written and oral forms; and integrating history, physical, and laboratory data with pathophysiologic principles. Attention to the economic, ethical, and sociological issues involved in patient care. Consists of immersive clinical experiences at Massachusetts General Hospital, leveraging extensive educational resources across inpatient clinical floors, ambulatory clinics, procedural/surgical suites, diagnostic testing areas, simulation learning lab, and didactic settings, followed by a focused experience in which students develop a proposal to solve an unmet need identified during their clinical experiences. Equivalent to combination of HST.201 and HST.202. Restricted to HST MEMP students.
P. Ankomah, C. Stultz, A. Puig, J. Ziperstein
HST.220 Introduction to the Care of Patients
Prereq: Permission of instructor
G (Spring)
1-0-2 units
Provides an introduction to the care of patients through opportunities to observe and participate in doctor-patient interaction in clinical settings and a longitudinal preceptorship experience with HST alumni physicians. Students are exposed to some of the practical realities of providing patient care. Topics include basic interviewing; issues of ethics, bias, and confidentiality; and other aspects of the doctor-patient relationship. The introductory session is held at HMS or Massachusetts General Hospital and the preceptorships are at several Harvard hospitals in Boston. Requirements include attendance at the introductory session and meetings scheduled with the preceptor.
N. Tetteh
HST.240 Translational Medicine Preceptorship
Prereq: HST.035
G (Fall, Spring, Summer)
0-12-0 units
Individually designed preceptorship joins together scientific research and clinical medicine. Students devote approximately half of their time to clinical experiences, and the remaining part to scholarly work in basic or clinical science. The two might run concomitantly or in series. Follow a clinical preceptor's daily activity, including aspects of patient care, attending rounds, conferences, and seminars. Research involves formal investigation of a focused and directed issue related to selected clinical area. Final paper required. Limited to students in the GEMS Program.
E. Edelman
HST.420[J] Principles and Practice of Assistive Technology
Same subject as 2.78[J], 6.4530[J]
Prereq: Permission of instructor
U (Fall)
Not offered regularly; consult department
2-4-6 units
See description under subject 6.4530[J]. Enrollment may be limited.
R. C. Miller, J. E. Greenberg, J. J. Leonard
HST.431[J] Infections and Inequalities: Interdisciplinary Perspectives on Global Health
Same subject as 11.134[J]
Prereq: None
U (Spring)
Not offered regularly; consult department
3-0-9 units. HASS-S
See description under subject 11.134[J]. Limited to 25.
E. James, A. Chakraborty
HST.434 Evolution of an Epidemic (Study Abroad)
Prereq: None
Acad Year 2024-2025: Not offered
Acad Year 2025-2026: U (IAP)
3-0-1 units
Examines the medical, scientific, public health and policy responses to a new disease, by focusing on the evolution of the AIDS epidemic. Begins with a review of how this new disease was first detected in the US and Africa, followed by the scientific basis as to how HIV causes profound dysfunction of the body's immune defense mechanisms, the rational development of drugs, the challenge of an HIV vaccine, and how public health and policy decisions have influenced the course of the global epidemic. Class conducted in Johannesburg Durban, South Africa. Open to all majors. Limited to 20. Application required; see class website for eligibility details.
H. Heller, B. Walker
HST.438[J] Viruses, Pandemics, and Immunity
Same subject as 5.002[J], 10.380[J]
Subject meets with 5.003[J], 8.245[J], 10.382[J], HST.439[J]
Prereq: None
U (Spring)
Not offered regularly; consult department
2-0-1 units
Covers the history of infectious diseases, basics of virology, immunology, and epidemiology, and ways in which diagnostic tests, vaccines, and antiviral therapies are currently designed and manufactured. Examines the origins of inequities in infection rates in society, and issues pertinent to vaccine safety. Final project explores how to create a more pandemic-resilient world. Subject can count toward the 6-unit discovery-focused credit limit for first-year students. Preference to first-year students; all others should take HST.439[J].
A. Chakraborty
HST.439[J] Viruses, Pandemics, and Immunity
Same subject as 5.003[J], 8.245[J], 10.382[J]
Subject meets with 5.002[J], 10.380[J], HST.438[J]
Prereq: None
U (Spring)
Not offered regularly; consult department
2-0-1 units
Covers the history of infectious diseases, basics of virology, immunology, and epidemiology, and ways in which diagnostic tests, vaccines, and antiviral therapies are currently designed and manufactured. Examines the origins of inequities in infection rates in society, and issues pertinent to vaccine safety. Final project explores how to create a more pandemic-resilient world. HST.438[J] intended for first-year students; all others should take HST.439[J].
A. Chakraborty
HST.450[J] Biological Physics
Same subject as 8.593[J]
Prereq: 8.044 recommended but not necessary
G (Spring)
Not offered regularly; consult department
4-0-8 units
See description under subject 8.593[J].
G. Benedek
HST.452[J] Statistical Physics in Biology
Same subject as 8.592[J]
Prereq: 8.333 or permission of instructor
Acad Year 2024-2025: G (Fall)
Acad Year 2025-2026: Not offered
3-0-9 units
A survey of problems at the interface of statistical physics and modern biology: bioinformatic methods for extracting information content of DNA; gene finding, sequence comparison, phylogenetic trees. Physical interactions responsible for structure of biopolymers; DNA double helix, secondary structure of RNA, elements of protein folding. Considerations of force, motion, and packaging; protein motors, membranes. Collective behavior of biological elements; cellular networks, neural networks, and evolution.
M. Kardar, L. Mirny
HST.460[J] Statistics for Neuroscience Research
Same subject as 9.073[J]
Prereq: Permission of instructor
Acad Year 2024-2025: Not offered
Acad Year 2025-2026: G (Spring)
3-0-9 units
See description under subject 9.073[J].
E. N. Brown
HST.482[J] Biomedical Signal and Image Processing
Same subject as 6.8801[J]
Subject meets with 6.8800[J], 16.456[J], HST.582[J]
Prereq: (6.3700 or permission of instructor) and (2.004, 6.3000, 16.002, or 18.085)
U (Spring)
3-1-8 units
See description under subject 6.8801[J].
J. Greenberg, E. Adalsteinsson, W. Wells
HST.500 Frontiers in (Bio)Medical Engineering and Physics
Prereq: None
G (Spring)
3-0-9 units
Provides a framework for mapping research topics at the intersection of medicine and engineering/physics in the Harvard-MIT community and covers the different research areas in MEMP (for example, regenerative biomedical technologies, biomedical imaging and biooptics). Lectures provide fundamental concepts and consider what's hot, and why, in each area. Training in scientific proposal writing (thesis proposals, fellowship applications, or research grant applications) through writing workshops. Topics include how to structure a novel research project, how to position research within the scientific community, how to present preliminary data effectively, and how to give and respond to peer reviews.
S. Bhatia, D. Anderson, S. Jhaveri
HST.504[J] Topics in Computational Molecular Biology
Same subject as 18.418[J]
Prereq: 6.8701, 18.417, or permission of instructor
G (Fall)
3-0-9 units
Can be repeated for credit.
See description under subject 18.418[J].
B. Berger
HST.506[J] Computational Systems Biology: Deep Learning in the Life Sciences
Same subject as 6.8710[J]
Subject meets with 6.8711[J], 20.390[J], 20.490
Prereq: Biology (GIR) and (6.3700 or 18.600)
Acad Year 2024-2025: G (Spring)
Acad Year 2025-2026: Not offered
3-0-9 units
See description under subject 6.8710[J].
D. K. Gifford
HST.507[J] Advanced Computational Biology: Genomes, Networks, Evolution
Same subject as 6.8700[J]
Subject meets with 6.8701
Prereq: (Biology (GIR), 6.1210, and 6.3700) or permission of instructor
G (Fall)
4-0-8 units
See description under subject 6.8700[J].
M. Kellis
HST.508[J] Evolutionary and Quantitative Genomics
Same subject as 1.872[J]
Prereq: Permission of instructor
Acad Year 2024-2025: Not offered
Acad Year 2025-2026: G (Fall)
4-0-8 units
Develops deep quantitative understanding of basic forces of evolution, molecular evolution, genetic variations and their dynamics in populations, genetics of complex phenotypes, and genome-wide association studies. Applies these foundational concepts to cutting-edge studies in epigenetics, gene regulation and chromatin; cancer genomics and microbiomes. Modules consist of lectures, journal club discussions of high-impact publications, and guest lectures that provide clinical correlates. Homework assignments and final projects develop practical experience and understanding of genomic data from evolutionary principles.
L. Mirny, T. Lieberman
HST.515[J] Aerospace Biomedical and Life Support Engineering
Same subject as 16.423[J], IDS.337[J]
Prereq: 16.06, 16.400, or permission of instructor
Acad Year 2024-2025: Not offered
Acad Year 2025-2026: G (Spring)
3-0-9 units
See description under subject 16.423[J].
D. J. Newman
HST.518[J] Human Systems Engineering
Same subject as 16.453[J]
Subject meets with 16.400
Prereq: 6.3700, 16.09, or permission of instructor
G (Fall)
3-0-9 units
See description under subject 16.453[J].
L. A. Stirling
HST.522[J] Biomaterials: Tissue Interactions
Same subject as 2.79[J]
Prereq: (Biology (GIR), Chemistry (GIR), and Physics I (GIR)) or permission of instructor
G (Fall)
Not offered regularly; consult department
3-0-9 units
Principles of materials science and cell biology underlying the development and implementation of biomaterials for the fabrication of medical devices/implants, including artificial organs and matrices for tissue engineering and regenerative medicine. Employs a conceptual model, the "unit cell process for analysis of the mechanisms underlying wound healing and tissue remodeling following implantation of biomaterials/devices in various organs, including matrix synthesis, degradation, and contraction. Methodology of tissue and organ regeneration. Discusses methods for biomaterials surface characterization and analysis of protein adsorption on biomaterials. Design of implants and prostheses based on control of biomaterials-tissue interactions. Comparative analysis of intact, biodegradable, and bioreplaceable implants by reference to case studies. Criteria for restoration of physiological function for tissues and organs.
I. V. Yannas, M. Spector
HST.523[J] Cell-Matrix Mechanics
Same subject as 2.785[J]
Prereq: (Biology (GIR), Chemistry (GIR), and 2.001) or permission of instructor
G (Fall)
Not offered regularly; consult department
3-0-9 units
See description under subject 2.785[J].
I. V. Yannas, M. Spector
HST.524[J] Design of Medical Devices and Implants
Same subject as 2.782[J]
Prereq: (Biology (GIR), Chemistry (GIR), and Physics I (GIR)) or permission of instructor
G (Spring)
3-0-9 units
See description under subject 2.782[J].
I. V. Yannas, M. Spector
HST.525[J] Tumor Microenvironment and Immuno-Oncology: A Systems Biology Approach
Same subject as 10.548[J]
Prereq: None
Acad Year 2024-2025: G (Fall)
Acad Year 2025-2026: Not offered
2-0-4 units
Provides theoretical background to analyze and synthesize the most up-to-date findings from both laboratory and clinical investigations into solid tumor pathophysiology. Covers different topics centered on the critical role that the tumor microenvironment plays in the growth, invasion, metastasis and treatment of solid tumors. Develops a systems-level, quantitative understanding of angiogenesis, extracellular matrix, metastatic process, delivery of drugs and immune cells, and response to conventional and novel therapies, including immunotherapies. Discussions provide critical comments on the challenges and the future opportunities in research on cancer and in establishment of novel therapeutic approaches and biomarkers to guide treatment.
R. K. Jain, L. Munn
HST.526[J] Future Medicine: Drug Delivery, Therapeutics, and Diagnostics
Same subject as 10.643[J]
Subject meets with 10.443
Prereq: 5.12 or permission of instructor
G (Spring)
Not offered regularly; consult department
3-0-6 units
See description under subject 10.643[J]. Limited to 40.
D. G. Anderson
HST.531 Medical Physics of Proton Radiation Therapy
Prereq: None
Acad Year 2024-2025: Not offered
Acad Year 2025-2026: G (Spring)
2-0-4 units
Acceleration of protons for radiation therapy; introduction into advanced techniques such as laser acceleration and dielectric wall acceleration. Topics include the interactions of protons with the patient, Monte Carlo simulation, and dose calculation methods; biological aspects of proton therapy, relative biological effectiveness (RBE), and the role of contaminating neutrons; treatment planning and treatment optimization methods, and intensity-modulated proton therapy (IMPT); the effect of organ motion and its compensation by use of image-guided treatment techniques; general dosimetry and advanced in-vivo dosimetry methods, including PET/CT and prompt gamma measurements. Outlook into therapy with heavier ions. Includes practical demonstrations at the Proton Therapy Center of the Massachusetts General Hospital.
B. Winey, J. Schuemann
HST.533 Medical Imaging in Radiation Therapy
Prereq: 18.06
Acad Year 2024-2025: G (Spring)
Acad Year 2025-2026: Not offered
2-0-4 units
Introduces imaging concepts and applications used throughout radiation therapy workflows, including magnetic resonance imaging (MRI), positron emission tomography (PET), and computed tomography (CT). Advanced topics include proton imaging modalities, such as prompt gamma imaging and proton radiography/CT. Includes lectures regarding image reconstruction and image registration. Introduces students to open-source medical image computing software (3D Slicer, RTK, and Plastimatch). Includes imaging demonstrations at Massachusetts General Hospital.
B. Winey, J. Schuemann
HST.535[J] Tissue Engineering and Organ Regeneration
Same subject as 2.787[J]
Prereq: (Biology (GIR), Chemistry (GIR), and Physics I (GIR)) or permission of instructor
G (Fall)
3-0-9 units
Principles and practice of tissue engineering (TE) and organ regeneration (OR). Topics include factors that prevent the spontaneous regeneration of tissues/organs in the adult (following traumatic injury, surgical excision, disease, and aging), and molecular and cell-biological mechanisms that can be harnessed for induced regeneration. Presents the basic science of organ regeneration. Principles underlying engineering strategies for employing select biomaterial scaffolds, exogenous cells, soluble regulators, and physical stimuli, for the formation of tissue in vitro (TE) and regeneration of tissues/organs in vivo (OR). Describes the technologies for producing biomaterial scaffolds and for incorporating cells and regulatory molecules into workable devices. Examples of clinical successes and failures of regenerative devices are analyzed as case studies.
M. Spector, I. V. Yannas
HST.537[J] Fluids and Diseases
Same subject as 1.631[J], 2.250[J]
Subject meets with 1.063
Prereq: None
Acad Year 2024-2025: G (Spring)
Acad Year 2025-2026: Not offered
3-3-6 units
See description under subject 1.631[J].
L. Bourouiba
HST.538[J] Genomics and Evolution of Infectious Disease
Same subject as 1.881[J]
Subject meets with 1.088
Prereq: Biology (GIR) and (1.000 or 6.100B)
G (Spring)
3-0-9 units
See description under subject 1.881[J].
T. Lieberman
HST.539[J] Advances in Interdisciplinary Science in Human Health and Disease
Same subject as 5.64[J]
Prereq: 5.13, 5.601, 5.602, and (5.07[J] or 7.05)
G (Spring)
3-0-9 units
See description under subject 5.64[J].
A. Shalek, X. Wang
HST.540[J] Human Physiology
Same subject as 7.20[J]
Prereq: 7.05
Acad Year 2024-2025: Not offered
Acad Year 2025-2026: U (Fall)
5-0-7 units
See description under subject 7.20[J].
M. Krieger, O. Yilmaz
HST.541[J] Cellular Neurophysiology and Computing
Same subject as 2.794[J], 6.4812[J], 9.021[J], 20.470[J]
Subject meets with 2.791[J], 6.4810[J], 9.21[J], 20.370[J]
Prereq: (Physics II (GIR), 18.03, and (2.005, 6.2000, 6.3000, 10.301, or 20.110[J])) or permission of instructor
G (Spring)
5-2-5 units
See description under subject 6.4812[J].
J. Han, T. Heldt
HST.542[J] Quantitative and Clinical Physiology
Same subject as 2.792[J], 6.4820[J]
Subject meets with 2.796[J], 6.4822[J], 16.426[J]
Prereq: Physics II (GIR), 18.03, or permission of instructor
U (Fall)
4-2-6 units
See description under subject 6.4820[J].
T. Heldt, R. G. Mark
HST.552[J] Medical Device Design
Same subject as 2.75[J], 6.4861[J]
Subject meets with 2.750[J], 6.4860[J]
Prereq: 2.008, 6.2040, 6.2050, 6.2060, 22.071, or permission of instructor
G (Spring)
3-3-6 units
See description under subject 2.75[J]. Enrollment limited.
A. H. Slocum, E. Roche, N. C. Hanumara, G. Traverso, A. Pennes
HST.560[J] Radiation Biophysics
Same subject as 22.55[J]
Subject meets with 22.055
Prereq: Permission of instructor
Acad Year 2024-2025: Not offered
Acad Year 2025-2026: G (Fall)
3-0-9 units
See description under subject 22.55[J].
Staff
HST.562[J] Pioneering Technologies for Interrogating Complex Biological Systems
Same subject as 9.271[J], 10.562[J]
Prereq: None
G (Spring)
3-0-9 units
Introduces pioneering technologies in biology and medicine and discusses their underlying biological/molecular/engineering principles. Topics include emerging sample processing technologies, advanced optical imaging modalities, and next-gen molecular phenotyping techniques. Provides practical experience with optical microscopy and 3D phenotyping techniques. Limited to 15.
K. Chung
HST.563 Imaging Biophysics and Clinical Applications
Prereq: (8.03 and 18.03) or permission of instructor
G (Spring)
Not offered regularly; consult department
2-1-9 units
Introduction to the connections and distinctions among various imaging modalities (x-ray, optical, ultrasound, MRI, PET, SPECT, EEG), common goals of biomedical imaging, broadly defined target of biomedical imaging, and the current practical and economic landscape of biomedical imaging research. Emphasis on applications of imaging research. Final project consists of student groups writing mock grant applications for biomedical imaging research project, modeled after an exploratory National Institutes of Health (NIH) grant application.
C. Catana
HST.565 Medical Imaging Sciences and Applications
Prereq: None
G (Fall)
Not offered regularly; consult department
3-0-9 units
Covers biophysical, biomedical, mathematical and instrumentation basics of positron emission tomography (PET), x-ray and computed tomography (CT), magnetic resonance imaging (MRI), single photon emission tomography (SPECT), optical Imaging and ultrasound. Topics include particles and photon interactions, nuclear counting statistics, gamma cameras, and computed tomography as it pertains to SPECT and PET (PET-CT, PET-MR, time-of-flight PET), MR physics and various sequences, optical and ultrasound physics foundations for imaging. Discusses clinical applications of PET and MR in molecular imaging of the brain, the heart, cancer and the role of AI in medical imaging. Includes medical demonstration lectures of SPECT, PET-CT and PET-MR imaging at Massachusetts General Hospital. Considers the ways imaging techniques are rooted in physics, engineering, and mathematics, and their respective role in anatomic and physiologic/molecular imaging.
HST Faculty
HST.576[J] Topics in Neural Signal Processing
Same subject as 9.272[J]
Prereq: Permission of instructor
Acad Year 2024-2025: G (Spring)
Acad Year 2025-2026: Not offered
3-0-9 units
See description under subject 9.272[J].
E. N. Brown
HST.580[J] Data Acquisition and Image Reconstruction in MRI
Same subject as 6.8810[J]
Prereq: 6.3010
Acad Year 2024-2025: Not offered
Acad Year 2025-2026: G (Fall)
3-0-9 units
See description under subject 6.8810[J].
E. Adalsteinsson
HST.582[J] Biomedical Signal and Image Processing
Same subject as 6.8800[J], 16.456[J]
Subject meets with 6.8801[J], HST.482[J]
Prereq: (6.3700 and (2.004, 6.3000, 16.002, or 18.085)) or permission of instructor
G (Spring)
3-1-8 units
See description under subject 6.8800[J].
J. Greenberg, E. Adalsteinsson, W. Wells
HST.583[J] Functional Magnetic Resonance Imaging: Data Acquisition and Analysis
Same subject as 9.583[J]
Prereq: 18.05 and (18.06 or permission of instructor)
Acad Year 2024-2025: G (Fall)
Acad Year 2025-2026: Not offered
2-3-7 units
Provides background necessary for designing, conducting, and interpreting fMRI studies in the human brain. Covers in depth the physics of image encoding, mechanisms of anatomical and functional contrasts, the physiological basis of fMRI signals, cerebral hemodynamics, and neurovascular coupling. Also covers design methods for stimulus-, task-driven and resting-state experiments, as well as workflows for model-based and data-driven analysis methods for data. Instruction in brain structure analysis and surface- and region-based analyses. Laboratory sessions include data acquisition sessions at the 3 Tesla MRI scanner at MIT and the Connectom and 7 Tesla scanners at the MGH/HST Martinos Center, as well as hands-on data analysis workshops. Introductory or college-level neurobiology, physics, and signal processing are helpful.
J. Polimeni, A. Yendiki, J. Chen
HST.584[J] Magnetic Resonance Analytic, Biochemical, and Imaging Techniques
Same subject as 22.561[J]
Prereq: Permission of instructor
Acad Year 2024-2025: G (Spring)
Acad Year 2025-2026: Not offered
3-0-12 units
Introduction to basic NMR theory. Examples of biochemical data obtained using NMR summarized along with other related experiments. Detailed study of NMR imaging techniques includes discussions of basic cross-sectional image reconstruction, image contrast, flow and real-time imaging, and hardware design considerations. Exposure to laboratory NMR spectroscopic and imaging equipment included.
L. Wald, B. Bilgic
HST.590 Biomedical Engineering Seminar Series
Prereq: None
G (Fall, Spring)
1-0-0 units
Can be repeated for credit.
Seminars focused on the development of professional skills for biomedical engineers and scientists. Each term focuses on a different topic, resulting in a repeating cycle that covers biomedical and research ethics, business and entrepreneurship, global health and biomedical innovation, and health systems and policy. Includes guest lectures, case studies, interactive small group discussions, and role-playing simulations.
E. Lindemer, M. Cotler
HST.599 Research in Health Sciences and Technology
Prereq: Permission of instructor
G (Fall, IAP, Spring, Summer)
Units arranged [P/D/F]
Can be repeated for credit.
For students conducting pre-thesis research or lab rotations in HST, in cases where the assigned research is approved for academic credit by the department. Hours arranged with research advisor. Restricted to HST students.
Consult Faculty
HST.714[J] Introduction to Sound, Speech, and Hearing
Same subject as 9.016[J]
Prereq: (6.3000 and 8.03) or permission of instructor
G (Fall)
Not offered regularly; consult department
4-0-8 units
Introduces students to the acoustics, anatomy, physiology, and mechanics related to speech and hearing. Focuses on how humans generate and perceive speech. Topics related to speech, explored through applications and challenges involving acoustics, speech recognition, and speech disorders, include acoustic theory of speech production, basic digital speech processing, control mechanisms of speech production and basic elements of speech and voice perception. Topics related to hearing include acoustics and mechanics of the outer ear, middle ear, and cochlea, how pathologies affect their function, and methods for clinical diagnosis. Surgical treatments and medical devices such as hearing aids, bone conduction devices, and implants are also covered.
S. S. Ghosh, H. H. Nakajima, S. Puria
HST.716[J] Signal Processing by the Auditory System: Perception
Same subject as 6.8830[J]
Prereq: (6.3000 and (6.3700 or 6.3702)) or permission of instructor
G (Fall)
Not offered regularly; consult department
3-0-9 units
See description under subject 6.8830[J].
L. D. Braida
HST.723[J] Audition: Neural Mechanisms, Perception and Cognition
Same subject as 9.285[J]
Prereq: Permission of instructor
G (Spring)
6-0-6 units
Neural structures and mechanisms mediating the detection, localization and recognition of sounds. General principles are conveyed by theme discussions of auditory masking, sound localization, musical pitch, cochlear implants, cortical plasticity and auditory scene analysis. Follows Harvard FAS calendar.
J. McDermott, D. Polley, M. C. Brown
HST.728[J] Spoken Language Processing
Same subject as 6.8620[J]
Prereq: 6.3000 and 6.3900
Acad Year 2024-2025: Not offered
Acad Year 2025-2026: G (Spring)
3-1-8 units
See description under subject 6.8620[J].
J. R. Glass
HST.916[J] Case Studies and Strategies in Drug Discovery and Development
Same subject as 7.549[J], 15.137[J], 20.486[J]
Prereq: None
G (Spring)
Not offered regularly; consult department
2-0-4 units
See description under subject 20.486[J].
A. W. Wood
HST.918[J] Economics of Health Care Industries
Same subject as 15.141[J]
Prereq: None
G (Spring; first half of term)
3-0-3 units
Credit cannot also be received for 15.1411
See description under subject 15.141[J].
J. Doyle
HST.920[J] Principles and Practice of Drug Development
Same subject as 10.547[J], 15.136[J], IDS.620[J]
Prereq: Permission of instructor
G (Fall)
3-0-6 units
See description under subject 15.136[J].
S. Finkelstein, A. J. Sinskey, R. Rubin
HST.936 Global Health Informatics to Improve Quality of Care
Subject meets with HST.937, HST.938
Prereq: None
Acad Year 2024-2025: Not offered
Acad Year 2025-2026: G (Spring)
2-0-1 units
Addresses issues related to how health information systems can improve the quality of care in resource poor settings. Discusses key challenges and real problems; design paradigms and approaches; and system evaluation and the challenges of measuring impact. Weekly lectures led by internationally recognized experts in the field. Students taking HST.936, HST.937 and HST.938 attend common lectures; assignments and laboratory time differ. HST.936 has no laboratory.
L. G. Celi, H. S. Fraser, V. Nikore, K. Paik, M. Somai
HST.937 Global Health Informatics to Improve Quality of Care
Subject meets with HST.936, HST.938
Prereq: None
Acad Year 2024-2025: Not offered
Acad Year 2025-2026: G (Spring)
2-2-2 units
Addresses issues related to how health information systems can improve the quality of care in resource poor settings. Discusses key challenges and real problems; design paradigms and approaches; and system evaluation and the challenges of measuring impact. Weekly lectures led by internationally recognized experts in the field. Students taking HST.936, HST.937 and HST.938 attend common lectures; assignments and laboratory time differ. HST.936 has no laboratory.
L. G. Celi, H. S. Fraser, V. Nikore, K. Paik. M. Somai
HST.938 Global Health Informatics to Improve Quality of Care
Subject meets with HST.936, HST.937
Prereq: None
Acad Year 2024-2025: Not offered
Acad Year 2025-2026: G (Spring)
2-2-8 units
Addresses issues related to how health information systems can improve the quality of care in resource poor settings. Discusses key challenges and real problems; design paradigms and approaches; and system evaluation and the challenges of measuring impact. Weekly lectures led by internationally recognized experts in the field. Students taking HST.936, HST.937 and HST.938 attend common lectures; assignments and laboratory time differ. HST.936 has no laboratory.
L. G. Celi, H. S. Fraser, V. Nikore, K. Paik, M. Somai
HST.940[J] Bioinformatics: Principles, Methods and Applications
Same subject as 10.555[J]
Prereq: Permission of instructor
G (Spring)
Not offered regularly; consult department
3-0-9 units
See description under subject 10.555[J].
Gr. Stephanopoulos, I. Rigoutsos
HST.953[J] Clinical Data Learning, Visualization, and Deployments
Same subject as 6.8850[J]
Prereq: (6.7900 and 6.7930[J]) or permission of instructor
G (Fall)
3-0-9 units
Examines the practical considerations for operationalizing machine learning in healthcare settings, with a focus on robust, private, and fair modeling using real retrospective healthcare data. Explores the pre-modeling creation of dataset pipeline to the post-modeling "implementation science," which addresses how models are incorporated at the point of care. Students complete three homework assignments (one each in machine learning, visualization, and implementation), followed by a project proposal and presentation. Students gain experience in dataset creation and curation, machine learning training, visualization, and deployment considerations that target utility and clinical value. Students partner with computer scientists, engineers, social scientists, and clinicians to better appreciate the multidisciplinary nature of data science.
M. Ghassemi, L. A. Celi, N. McCague and E. Gottlieb
HST.956[J] Machine Learning for Healthcare
Same subject as 6.7930[J]
Prereq: 6.3900, 6.4100, 6.7810, 6.7900, 6.8611, or 9.520[J]
G (Spring)
4-0-8 units
See description under subject 6.7930[J]. Limited to 55.
D. Sontag, P. Szolovits
HST.962 Medical Product Development and Translational Biomedical Research
Prereq: Permission of instructor
G (Spring; second half of term)
1-0-3 units
Explores the translation of basic biomedical science into therapies. Topics span pharmaceutical, medical device, and diagnostics development. Exposes students to strategic assessment of clinical areas, product comparison, regulatory risk assessment by indication, and rational safety program design. Develops quantitative understanding of statistics and trial design.
M. Cima
HST.971[J] Strategic Decision Making in Life Science Ventures
Same subject as 15.363[J]
Prereq: None
G (Spring)
3-0-6 units
See description under subject 15.363[J].
J. Fleming, A. Zarur
HST.974 Innovating for Mission Impact in Medicine and Healthcare
Prereq: Permission of instructor
G (Fall, Spring)
3-0-9 units
Can be repeated for credit.
Through a mentored experience, and in conjunction with the MIT Catalyst program, participants develop and validate a small portfolio of research opportunities/proposals. Provides experience with critical professional skills (interfacing with diverse experts, research strategy, critically evaluating the landscape and potential to add value, proposal development, communication, etc.) that heightens the potential to have meaningful impact through their work and career. Restricted to MIT Catalyst Fellows.
M. Gray, B. Vakoc, T. Padera
HST.978[J] Healthcare Ventures
Same subject as 15.367[J]
Prereq: None
G (Spring)
3-0-9 units
Addresses healthcare entrepreneurship with an emphasis on startups bridging care re-design, digital health, medical devices, and new healthcare business models. Includes prominent speakers and experts from key domains across venture capital, medicine, pharma, med devices, regulatory, insurance, software, design thinking, entrepreneurship, including many alumni from the class sharing their journeys. Provides practical experiences in venture validation/creation through team-based work around themes. Illustrates best practices in identifying and validating health venture opportunities amid challenges of navigating healthcare complexity, team dynamics, and venture capital raising process. Intended for students from engineering, medicine, public health, and MBA programs. Video conference facilities provided to facilitate remote participation by Executive MBA and traveling students.
M. Gray, Z. Chu
HST.980 Emerging Problems in Infectious Diseases
Prereq: None
G (IAP)
1-0-2 units
Introduces contemporary challenges in preventing, detecting, diagnosing and treating emerging and newly emerging pathogens. Provides students with team-based opportunities to brainstorm, propose and present innovative solutions to such challenges. Expert lecturers discuss emerging problems in infectious diseases. Includes brainstorming sessions in which student teams identify problems in infectious diseases and propose innovative solutions. The teams then prepare and deliver short presentations, outlining identified problems and solutions.
J. J. Collins
HST.999 Practical Experience in Health Sciences and Technology
Prereq: None
G (Fall, IAP, Spring, Summer)
Units arranged [P/D/F]
Can be repeated for credit.
Required for HST PhD students to gain professional perspective in research experiences, academic experiences, or internships related to health sciences and technology. Professional perspective options include: internships (with industry, government, medicine or academia), industrial or medical colloquia or seminars, research collaboration with industry or government, and professional development for entry into academia or entrepreneurial engagement. For an internship experience, an offer of employment from a company or organization is required prior to enrollment. Upon completion of the activity, student must submit a letter from the employer describing the work accomplished, along with a substantive final report written by the student. Consult HST's Academic Office for details on procedures and restrictions.
J. Greenberg
HST.THG Graduate Thesis
Prereq: Permission of instructor
G (Fall, IAP, Spring, Summer)
Units arranged
Can be repeated for credit.
Program of research leading to the writing of a PhD or ScD thesis or an HST SM thesis; to be arranged by the student and an appropriate faculty advisor.
Faculty
HST.UR Undergraduate Research in Health Sciences and Technology
Prereq: None
U (Fall, IAP, Spring, Summer)
Units arranged [P/D/F]
Can be repeated for credit.
Extended participation in the work of a faculty member or research group. Research is arranged by mutual agreement between the student and a member of the faculty of the Harvard-MIT Program Health Sciences and Technology, and may continue over several terms. Registration requires submission of a written proposal to the MIT UROP, signed by the faculty advisor and approved by the department. A summary report must be submitted at the end of each term.
HST Faculty
HST.URG Undergraduate Research in Health Sciences and Technology
Prereq: None
U (Fall, IAP, Spring, Summer)
Units arranged
Can be repeated for credit.
Extended participation in the work of a faculty member or research group. Research is arranged by mutual agreement between the student and a member of the faculty of the Harvard-MIT Program in Health Sciences and Technology, and may continue over several terms. Registration requires submission of a written proposal to the MIT UROP Office; signed by the faculty advisor and approved by the department. A summary report must be submitted at the end of each term.
HST Faculty
HST.S16 Special Graduate Subject: Health Sciences and Technology
Prereq: None
G (Fall, IAP, Spring, Summer)
Not offered regularly; consult department
Units arranged [P/D/F]
Can be repeated for credit.
Opportunity for group study of advanced subjects related to Health Sciences and Technology not otherwise included in the curriculum. Offerings are initiated by IMES/HST faculty on an ad hoc basis subject to program approval. Prerequisites may vary by topic; consult faculty at time of offering.
IMES/HST Faculty
HST.S17 Special Graduate Subject: Health Sciences and Technology
Prereq: Permission of instructor
G (Fall, IAP, Spring, Summer)
Not offered regularly; consult department
Units arranged [P/D/F]
Can be repeated for credit.
Opportunity for group study of advanced subjects related to Health Sciences and Technology not otherwise included in the curriculum. Offerings are initiated by IMES/HST faculty on an ad hoc basis subject to program approval. Prerequisites may vary by topic; consult faculty at time of offering.
IMES/HST Faculty
HST.S18 Special Graduate Subject: Health Sciences and Technology
Prereq: Permission of instructor
G (Fall, IAP, Spring, Summer)
Not offered regularly; consult department
Units arranged
Can be repeated for credit.
Opportunity for group study of advanced subjects related to Health Sciences and Technology not otherwise included in the curriculum. Offerings are initiated by IMES/HST faculty on an ad hoc basis subject to program approval. Prerequisites may vary by topic; consult faculty at time of offering.
IMES/HST Faculty
HST.S19 Special Graduate Subject: Health Sciences and Technology
Prereq: Permission of instructor
G (Fall, IAP, Spring, Summer)
Not offered regularly; consult department
Units arranged
Can be repeated for credit.
Opportunity for group study of advanced subjects related to Health Sciences and Technology not otherwise included in the curriculum. Offerings are initiated by IMES/HST faculty on an ad hoc basis subject to program approval. Prerequisites may vary by topic; consult faculty at time of offering.
IMES/HST Faculty
HST.S46 Special Undergraduate Subject: Health Sciences and Technology
Prereq: Permission of instructor
U (Fall, IAP, Spring, Summer)
Not offered regularly; consult department
Units arranged [P/D/F]
Can be repeated for credit.
Group study of subjects related to health sciences and technology not otherwise included in the curriculum. Prerequisites may vary by topic; consult faculty at time of offering.
IMES/HST Faculty
HST.S47 Special Undergraduate Subject: Health Sciences and Technology
Prereq: Permission of instructor
U (Fall, IAP, Spring, Summer)
Not offered regularly; consult department
Units arranged [P/D/F]
Can be repeated for credit.
Group study of subjects related to health sciences and technology not otherwise included in the curriculum. Prerequisites may vary by topic; consult faculty at time of offering.
IMES/HST Faculty
HST.S48 Special Undergraduate Subject: Health Sciences and Technology
Prereq: Permission of instructor
U (Fall, IAP, Spring, Summer)
Not offered regularly; consult department
Units arranged
Can be repeated for credit.
Group study of subjects related to health sciences and technology not otherwise included in the curriculum. Prerequisites may vary by topic; consult faculty at time of offering.
IMES/HST Faculty
HST.S49 Special Undergraduate Subject: Health Sciences and Technology
Prereq: Permission of instructor
U (Fall, IAP, Spring, Summer)
Not offered regularly; consult department
Units arranged
Can be repeated for credit.
Group study of subjects related to health sciences and technology not otherwise included in the curriculum. Prerequisites may vary by topic; consult faculty at time of offering.
IMES/HST Faculty
HST.S56 Special Graduate Subject: Medical Engineering and Medical Physics
Prereq: Permission of instructor
G (Fall)
Units arranged [P/D/F]
Can be repeated for credit.
Opportunity for group study of advanced subjects related to the Medical Engineering and Medical Physics Program not otherwise included in the curriculum. Offerings are initiated by IMES/HST faculty on an ad hoc basis subject to program approval. Prerequisites may vary by topic; consult faculty at time of offering.
IMES/HST Faculty
HST.S57 Special Graduate Subject: Medical Engineering and Medical Physics
Prereq: Permission of instructor
G (Fall, IAP, Spring, Summer)
Not offered regularly; consult department
Units arranged [P/D/F]
Can be repeated for credit.
Opportunity for group study of advanced subjects related to the Medical Engineering and Medical Physics Program not otherwise included in the curriculum. Offerings are initiated by IMES/HST faculty on an ad hoc basis subject to program approval. Prerequisites may vary by topic; consult faculty at time of offering.
IMES/HST Faculty
HST.S58 Special Subject: Medical Engineering and Medical Physics
Prereq: Permission of instructor
G (Fall, IAP, Spring, Summer)
Not offered regularly; consult department
Units arranged
Can be repeated for credit.
Opportunity for group study of advanced subjects related to the Medical Engineering and Medical Physics Program not otherwise included in the curriculum. Offerings are initiated by HST faculty on an ad hoc basis subject to program approval. Prerequisites may vary by topic; consult faculty at time of offering.
HST Faculty
HST.S59 Special Graduate Subject: Medical Engineering and Medical Physics
Prereq: Permission of instructor
G (Fall, IAP, Spring, Summer)
Not offered regularly; consult department
Units arranged
Can be repeated for credit.
Opportunity for group study of advanced subjects related to the Medical Engineering and Medical Physics Program not otherwise included in the curriculum. Offerings are initiated by IMES/HST faculty on an ad hoc basis subject to program approval. Prerequisites may vary by topic; consult faculty at time of offering.
IMES/HST Faculty
HST.S78 Special Subject: Speech and Hearing Sciences
Prereq: Permission of instructor
G (Fall, IAP, Spring)
Not offered regularly; consult department
Units arranged
Can be repeated for credit.
Opportunity for group study of advanced subjects related to the Speech and Hearing Sciences not otherwise included in the curriculum. Offerings initiated by members of the SHS faculty on an ad hoc basis subject to program approval. Prerequisites may vary by topic; consult faculty at time of offering.
P. Cariani
HST.S96 Special Graduate Subject: Biomedical Entrepreneurship
Prereq: Permission of instructor
G (Fall, IAP, Spring, Summer)
Not offered regularly; consult department
Units arranged [P/D/F]
Can be repeated for credit.
Opportunity for group study of advanced subjects relating to biomedical entrepreneurship not otherwise included in the curriculum. Offerings are initiated by HST/IMES faculty on an ad hoc basis subject to program approval. Prerequisites may vary by topic. Consult faculty at time of offering.
HST/IMES Faculty
HST.S97 Special Graduate Subject: Biomedical Entrepreneurship
Prereq: Permission of instructor
G (Fall, IAP, Spring, Summer)
Not offered regularly; consult department
Units arranged [P/D/F]
Can be repeated for credit.
Opportunity for group study of advanced subjects relating to biomedical entrepreneurship not otherwise included in the curriculum. Offerings are initiated by HST faculty on an ad hoc basis subject to program approval. Prerequisites may vary by topic. Consult faculty at time of offering.
HST Faculty
HST.S98 Special Graduate Subject: Biomedical Entrepreneurship
Prereq: Permission of instructor
G (Fall, Spring)
Not offered regularly; consult department
Units arranged
Can be repeated for credit.
Opportunity for group study of advanced subjects relating to biomedical entrepreneurship not otherwise included in the curriculum. Offerings are initiated by HST faculty on an ad hoc basis subject to program approval. Prerequisites may vary by topic. Consult faculty at time of offering.
M. Gray, F. Murray
HST.S99 Special Graduate Subject: Biomedical Entrepreneurship
Prereq: Permission of instructor
G (Spring)
Units arranged
Can be repeated for credit.
Opportunity for group study of advanced subjects relating to biomedical entrepreneurship not otherwise included in the curriculum. Offerings are initiated by HST faculty on an ad hoc basis subject to program approval. Prerequisites may vary by topic. Consult faculty at time of offering.
HST/IMES Faculty
